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Introduction

A panoramic view from a ridge located between Segla and Hesten mountain summits in the island of Senja, Troms, Norway in 2014
A panoramic view from a ridge located between Segla and Hesten mountain summits in the island of Senja, Troms, Norway in 2014

Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of at least one cell that processes hereditary information encoded in genes, which can be transmitted to future generations. Another major theme is evolution, which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move, grow, and reproduce. Finally, all organisms are able to regulate their own internal environments.

Biologists are able to study life at multiple levels of organization, from the molecular biology of a cell to the anatomy and physiology of plants and animals, and evolution of populations. Hence, there are multiple subdisciplines within biology, each defined by the nature of their research questions and the tools that they use. Like other scientists, biologists use the scientific method to make observations, pose questions, generate hypotheses, perform experiments, and form conclusions about the world around them.

Life on Earth, which emerged more than 3.7 billion years ago, is immensely diverse. Biologists have sought to study and classify the various forms of life, from prokaryotic organisms such as archaea and bacteria to eukaryotic organisms such as protists, fungi, plants, and animals. These various organisms contribute to the biodiversity of an ecosystem, where they play specialized roles in the cycling of nutrients and energy through their biophysical environment. (Full article...)

In multicellular organisms, stem cells are undifferentiated or partially differentiated cells that can change into various types of cells and proliferate indefinitely to produce more of the same stem cell. They are the earliest type of cell in a cell lineage. They are found in both embryonic and adult organisms, but they have slightly different properties in each. They are usually distinguished from progenitor cells, which cannot divide indefinitely, and precursor or blast cells, which are usually committed to differentiating into one cell type.

In mammals, roughly 50 to 150 cells make up the inner cell mass during the blastocyst stage of embryonic development, around days 5–14. These have stem-cell capability. In vivo, they eventually differentiate into all of the body's cell types (making them pluripotent). This process starts with the differentiation into the three germ layers – the ectoderm, mesoderm and endoderm – at the gastrulation stage. However, when they are isolated and cultured in vitro, they can be kept in the stem-cell stage and are known as embryonic stem cells (ESCs). (Full article...)

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Chimaeras are cartilaginous fish in the order Chimaeriformes, known informally as ghost sharks, ratfish, spookfish, or rabbitfishes. They grow up to 150 cm (4.9 ft) in length, and have elongated, soft bodies, with a bulky head and a single gill-opening. For defense, most chimaeras have a venomous spine located in front of the dorsal fin. At one time a "diverse and abundant" group (based on the fossil record), their closest living relatives are sharks, though in evolutionary terms they branched off from sharks nearly 400 million years ago and have remained isolated ever since, typically confined to deep water.

Major topics

History History of biology | timeline of biology and organic chemistry | history of ecology | history of evolutionary thought | history of geology | history of model organisms | history of molecular biology | history of paleontology
Overview Biology | science | life | properties (adaptationenergy processinggrowthorderregulationreproduction, and response to environment) | hierarchy of life (atommoleculeorganellecelltissueorganorgan systemorganismpopulationcommunityecosystembiosphere) | reductionistic | emergent property | mechanistic | scientific method | theory | law | peer review | biology journals
Chemical basis Matter | elements | compounds | atoms | molecules | chemical bonds | carbon | organic compounds | macromolecules | carbohydrate | protein | protein structure | protein folding | lipid | DNA | RNA
Cells Prokaryote | eukaryote | cell wall | cell membrane | cytoskeleton | mitochondrion | chloroplast | nucleus | endoplasmic reticulum | Golgi apparatus | cell cycle | mitosis | metabolism | cell signaling | protein targeting | metabolism | enzyme | glycolysis | citric acid cycle | electron transport chain | oxidative phosphorylation |photosynthesis |meiosis  | mitosis
Genetics (Intro) Classical genetics | mendelian inheritance | gene | phenotype | genotype | ploidy | alternation of generations | molecular genetics | gene expression | gene regulation | genome | karyotype | DNA replication | transcription | translation | recombination | chromosome | epigenetics | splicing | mutation | genetic fingerprint | chromatin | ecological genetics | population genetics | quantitative genetics
Evolution (Intro)  | omne vivum ex ovo | Natural selection | genetic drift | sexual selection | speciation | mutation | gene flow | evolution of sex | biogeography | cladistics | species | extinction | tree of life | phylogenies | three-domain system
Diversity Bacteria | archaea | plants | angiosperms | fungi | protists | Animals | deuterostome | insects | molluscs | nematodes | parasitism | Primate | mammal | vertebrate | craniata | chordate | viruses
Plant form and function Epidermis | flower | ground tissue  | leaf | phloem | plant stem | root | shoot | vascular plant | vascular tissue | xylem
Animal form and function Tissues | fertilization | embryogenesis | gastrulation | neurulation | organogenesis | differentiation | morphogenesis | metamorphosis | ontogeny  | Development | senescence  | reproduction | oogenesis | spermatogenesis
Ecology Ecosystem | biomass | food chain | indicator species | habitat | species distribution | Gaia theory | metapopulation  | life cycle | Life history | altricial - precocial | sex ratio | altruism | cooperation - foraging | learning | parental care | sexual conflict | territoriality | biosphere | climate change | conservation | biodiversity | nature reserve | edge effect | allee effect | corridor | fragmentation | pollution | invasive species | in situ - ex situ | seedbank
Research methods Laboratory techniques | Genetic engineering | transformation | gel electrophoresis | chromatography | centrifugation | cell culture | DNA sequencing | DNA microarray | green fluorescent protein | vector | enzyme assay | protein purification | Western blot | Northern blot | Southern blot | restriction enzyme | polymerase chain reaction | two-hybrid screening | in vivo - in vitro - in silico | Field techniques | Belt transect | mark and recapture | species discovery curve
Branches Anatomy | biotechnology | botany | cell biology | ecology | evolutionary biology | genetics | marine biology | microbiology | molecular biology | mycology | neuroscience | paleontology | phycology | physiology | protistology | virology | zoology
Awards Nobel Prize in Physiology or Medicine
See also Template:History of biology

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Tawfiq Canaan (Arabic: توفيق كنعان) (24 September 1882 – 15 January 1964) was a pioneering Palestinian physician, medical researcher, ethnographer, and Palestinian nationalist. Born in Beit Jala during the rule of the Ottoman Empire, he served as a medical officer in the Ottoman army during World War I.

During British rule, he served as the first President of the Palestine Arab Medical Association founded in 1944, and as the director of several Jerusalem area hospitals before, during, and after the 1948 war. Over the course of his medical career, he authored more than thirty-seven studies on topics including tropical medicine, bacteriology, malaria, tuberculosis, and health conditions in Palestine, and contributed to research that led to a cure for leprosy. (Full article...)

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